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Oleksandr Usyk of Ukraine is congratulated after his unanimous decision victory over Anthony Joshua of Britain in their WBA (Super), WBO and IBF boxing title bout at the Tottenham Hotspur Stadium in London, Saturday, Sept. 25, 2021. (AP Photo/Frank Augstein)

Slumped helplessly on the ropes, Anthony Joshua greeted the final bell — and what he knew was the end of his second reign as world heavyweight champion — with a wry smile.

Oleksandr Usyk knew it, too, dropping to his knees and looking to the sky.

In just his third fight since stepping up from the cruiserweight division, Usyk delivered a clinic in movement and fast punching to claim a unanimous points win over Joshua in front of 66,267 stunned fans inside Tottenham Hotspur Stadium in London on Saturday.

Usyk became the third man, after Evander Holyfield and David Haye, to win world titles at cruiserweight then heavyweight. Six years after Wladimir Klitschko’s long heavyweight reign ended, Ukraine has another champion in boxing’s marquee division.

Oleksandr Usyk of Ukraine, left, throws a punch at Anthony Joshua of Britain during their WBA (Super), WBO and IBF boxing title bout at the Tottenham Hotspur Stadium in London, Saturday, Sept. 25, 2021. (AP Photo/Frank Augstein)

“The fight went exactly as I expected it to go,” Usyk said through a translator. “There were a couple of moments when Anthony pushed me hard but nothing special.”

After the judges scored the fight 117-112, 116-112 and 115-113 in favor of the 34-year-old Usyk, Joshua — sporting a closed right eye — walked over to Usyk’s corner and congratulated the Ukrainian’s team.

Dominated by a smaller, lighter opponent, Joshua lost his WBA, WBO and IBF titles for a second time — more than two years after a defeat to Andy Ruiz Jr. in New York that he immediately avenged in Saudi Arabia.

Joshua will have the chance to do the same thing — there is a rematch clause in the deal with Usyk — in a bid to keep alive his ultimate goal, to become undisputed champion.

That looks fanciful now, though. Usyk was comfortably the more assured fighter, his reflexes drawing gasps from the pro-Joshua crowd and his lightning-quick punch off his left hand repeatedly connecting.

Oleksandr Usyk of Ukraine hits Anthony Joshua of Britain during their WBA (Super), WBO and IBF boxing title bout at the Tottenham Hotspur Stadium in London, Saturday, Sept. 25, 2021. (AP Photo/Frank Augstein)

Oleksandr Usyk of Ukraine celebrates after his unanimous decision victory over Anthony Joshua of Britain in their WBA (Super), WBO and IBF boxing title bout at the Tottenham Hotspur Stadium in London, Saturday, Sept. 25, 2021. (AP Photo/Frank Augstein)

Joshua just didn’t see the punches coming, like in the third round when his head was rocked back by a straight left before the Briton wobbled after a left hook flush on the jaw.

Like in the ninth, when another left from Usyk caught Joshua, who nodded to the challenger as the bell went.

And especially in a sensational final round when the disparity between the fighters was at its biggest as Usyk unleashed a flurry of punches — from his left and right — that saw Joshua staggering backward onto the ropes. Indeed, he was saved by the bell.

Joshua’s promoter, Eddie Hearn, said Joshua couldn’t see out of his right eye from the ninth round.

“I had no objective to knock him out because my corner pushed me not to do that,” Usyk said. “At the start, I hit him and tried to knock him out but my trainer said, ‘Just stop and do your job.’”

Oleksandr Usyk of Ukraine, left, throws a punch at Anthony Joshua of Britain during their WBA (Super), WBO and IBF boxing title bout at the Tottenham Hotspur Stadium in London, Saturday, Sept. 25, 2021. (AP Photo/Frank Augstein)Oleksandr Usyk of Ukraine holds up the Ukrainian flag after his unanimous decision victory over Anthony Joshua of Britain in their WBA (Super), WBO and IBF boxing title bout at the Tottenham Hotspur Stadium in London, Saturday, Sept. 25, 2021. (AP Photo/Frank Augstein)

Usyk stepped up to heavyweight in 2019 after unifying all the cruiserweight belts, and now has 19 straight wins as a professional.

His energy levels were remarkable, as shown in the final round and then after the result was announced as Usyk — an eccentric — entertained the crowd with an acrobatic dance inside the ring.

Joshua fell to the second loss of his 25-fight pro career, this time to a man who followed him — a day later — as Olympic champion at the London Games in 2012. His career is at a crossroads again.Oleksandr Usyk of Ukraine, left, throws a punch at Anthony Joshua of Britain during their WBA (Super), WBO and IBF boxing title bout at the Tottenham Hotspur Stadium in London, Saturday, Sept. 25, 2021. (AP Photo/Frank Augstein)

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